Explanation:

Let's analyze step by step:

- Expression: L = K = P = N = J @ U = A % R

- We need to replace (@) and (%) to ensure:

1. L = U

2. R > J

Checking all options:

Option 1: @ = =, % = =

- J = U (J can be less than or equal to U)

- A = R (A equals R)

- Hard to ensure R > J always as R=A, and A = U = J, possible contradictions.

Option 2: @ = =, % = =

- J = U

- A = R

- R might not be greater than J definitely, if all are equal.

Option 3: @ = >, % = =

- J > U

- A = R

- Violates L = U because if J > U and L is behind J, L may not be = U.

Option 4: @ = =, % = <

- J = U

- A < R

- L = K = P = N = J (= U) ? So, L = U (true)

- R > A = U (=J), so R > J (true)

- Both conditions are satisfied.

Option 5: @ = =, % = <

- J = U

- A < R

- J could be more than U, L = U not always certain.

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Conclusion:

Correct Option:

Option 4: =, <

- This ensures both L = U and R > J are definitely true.