Explanation:

Burrowing animals and termites: Termites, especially, are known to play a crucial role in savannah ecosystems by breaking down tough plant fibers, recycling dead plant matter, and enriching the soil. Their activities might not directly prevent forest development, but they play a role in shaping the savannah ecosystem.

Fire: Fires are a common occurrence in savannah ecosystems. They are an essential factor in maintaining the grassland by preventing the establishment and spread of trees. Fires can be naturally caused (like by lightning) or be a result of human activities.

Grazing herbivores: Savannahs are home to a large number of grazing animals like zebras, antelopes, and elephants. These herbivores feed on grasses and shrubs, preventing them from becoming too dominant and thus playing a role in shaping the landscape.

Seasonal rainfall: Savannahs typically have a wet season and a dry season. The prolonged dry season can make it challenging for many tree species to survive and establish, giving an advantage to grasses that are adapted to such conditions.

Soil properties: While soil properties can influence the type of vegetation in an area, it's not the primary limiting factor for forest development in savannahs. However, certain savannah regions might have soils that are less suitable for forest growth.

From the above analysis, the main conditions that limit forest development in savannahs are Fire (2), Grazing herbivores (3), and Seasonal rainfall (4).

Hence, the correct answer is option 3: 2, 3, and 4.